Land snails of Bozcaada island, Turkey with anatomical and distributional notes for several species
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Section of Mollusks, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, U.S.A.
Submission date: 2022-01-30
Final revision date: 2022-02-17
Acceptance date: 2022-02-19
Online publication date: 2022-03-05
Publication date: 2022-03-05
Corresponding author
Aydin Örstan   

Section of Mollusks, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, 15213, U.S.A.
Folia Malacol. 2022;30(1):36-46
A survey of the land snails of Bozcaada, a small island in the Aegean Sea near the northwest coast of Turkey, found 30 species. Mediterranea depressa (Sterki) was recorded for the first time in Turkey. Five species in the family Enidae were found on the island; the most unexpected one was Chondrus tournefortianus (Férussac) outside of its previously known range. The conchological and anatomical characteristics of the specimens identified as Meijeriella cf. raynevaliana (Bourguignat) are given and the associated taxonomic issues are briefly discussed. Anatomies of Monacha claustralis (Rossmässler), Orculella critica (Pfeiffer) and Oxychilus cyprius (Pfeiffer) are also presented. As expected, the land snail faunas of Bozcaada, the nearby islands of Gökçeada and Limnos and the mainland Turkey share some species. Some of the differences in the species compositions of these areas may be explained as results of human-mediated introductions.
Acıoğlu Y. 2015. Çanakkale’deki Osmanlı Kaleleri. Çanakkale Araştırmaları Türk Yıllığı 13: 93–122.
Anonymous 1916. Mediterranean Pilot. Vol. 4. Government Printing Office, Washington, pp. 383–384.
Bank R. A. 1985. Eine neue Enide von der griechischen Insel Mytilini (Gastropoda: Pupillacea). Heldia 1: 41–44.
Bank R. A., Gittenberger E., Neubert E. 2013. Radiation of an eastern Mediterranean landsnail genus: revision of the taxa belonging to Metafruticicola von Ihering 1892 (Gastropoda, Pulmonata: Hygromiidae). Archiv für Molluskenkunde 142: 67–136.
Bourguignat J. R. 1855. Aménités malacologiques. Revue et Magasin de Zoologie pure et appliquée 7: 556–574.
Clark P. U., Mix A. C. 2002. Ice sheets and sea level of the Last Glacial Maximum. Quaternary Science Reviews 21: 1–7.
Erguvanli K. 1955. Etude geologique de l’ile de Bozcaada. Bulletin de la Société Géologique de France. S6-V: 399–401.
Forcart L. 1940. Monographie der türkischen Enidae (Moll., Pulm.). Verhandlungen der Naturforschenden Gesellschaft in Basel 51: 106–263.
Georgiev D., Stoycheva S. 2010. Notes on the ecology and species diversity of the inland molluscs of Samothraki Island (North-Eastern Greece). North-Western Journal of Zoology 6: 71–78.
Gittenberger E., Hausdorf B. 2004. The Orculella species of the South Aegean island arc, a neglected radiation (Gastropoda, Pulmonata, Orculidae). Basteria 68: 93–124.
Gümüş B. A., Neubert E. 2012. New taxa of terrestrial molluscs from Turkey (Gastropoda, Pristilomatidae, Enidae, Hygromiidae, Helicidae). ZooKeys 171: 17–37.
Hausdorf B. 1996. Die Orculidae Asiens (Gastropoda: Stylommatophora). Archiv für Molluskenkunde 125: 1–86.
Hausdorf B. 2000. The genus Monacha in Turkey. Archiv für Molluskenkunde 128: 61–151.
Hausdorf B. 2001. The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627–1638.
Kerney M. P., Cameron R. A. D. 1979. A field guide to the land snails of Britain and North-west Europe. Collins, London.
NCEI (National Centers for Environmental Information) 2022. Bathymetric Data Viewer. Available online at (accessed 23 January 2022).
MolluscaBase 2021. MolluscaBase. Available online at (accessed 31 December 2021).
Neubert E. 2014. Revision of Helix Linnaeus, 1758 in its eastern Mediterranean distribution area, and reassignment of Helix godetiana Kobelt, 1878 to Maltzanella Hesse, 1917 (Gastropoda, Pulmonata, Helicidae). Contributions to Natural History 26: 1–200.
Nordsieck H. 1999. Annotated check-list of the species of the Albinaria-Isabellaria group (Gastropoda: Stylommatophora: Clausiliidae). Mitteilungen der Deutschen Malakozoologischen Gesellschaft 62/63: 1–21.
Örstan A. 2001. A preliminary survey of Albinaria populations around Kuşadasi Bay, Turkey. Triton 4: 42–44.
Örstan A. 2014. A survey of the land snails of the Karaburun peninsula, Turkey. Folia Malacologica 22: 201–207.
Örstan A., Yildirim M. Z. 2004. Borlumastus gen. nov. (Gastropoda, Pulmonata, Enidae), a new land snail genus from Turkey. Basteria 68: 125–129.
Pieńkowska J. R., Proćków M., Górka M., Lesicki A. 2018. Distribution of Monacha claustralis (Rossmässler, 1834) and M. cartusiana (O. F. Müller, 1774) (Eupulmonata: Hygromiidae) in central European and Balkan countries: new data. Folia Malacologica 26: 103–120.
Pintér L. 1972. Die Gattung Vitrea Fitzinger, 1833 in den Balkanländern (Gastropoda: Zonitidae). Annales Zoologici 29: 210–315.
Reischütz P. L. 1983. Ein Beitrag zur Molluskenfauna der Insel Thasos. Annalen des Naturhistorischen Museums in Wien 85B: 133–146.
Reischütz P. L. 1986. Beiträge zur Molluskenfauna der Ägäischen Inseln. Malakologische Abhandlungen 11: 93–101.
Reischütz P. L. 1988. Beiträge zur Molluskenfauna Thrakiens und Ostmakedoniens, II. Annalen des Naturhistorischen Museums in Wien 90B: 341–356.
Riedel A. 1991. Zonitidae und Daudebardiidae von Zypern (Gastropoda, Stylommatophora). Malakologische Ab­handlungen 15: 101–110.
Riedel A. 1992. The Zonitidae (sensu lato) (Gastropoda, Pulmonata) of Greece. Fauna Graeciae. Vol. 5. Athens.
Riedel A. 1995. Zonitidae sensu lato (Gastropoda, Stylommatophora) der Türkei. Übersicht der Arten. Fragmenta Faunistica 38: 1–86.
Schileyko A. A. 1998. Treatise on recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidae, Hypselo­stomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica, Supplement 2: 129–261.
Schütt H. 2010. Turkish land snails. Verlag Natur & Wissenschaft, Solingen.
Triantis K. A., Mylonas M., Weiser M. D., Lika K., Vardinoyannis K. 2005. Species richness, environmental heterogeneity and area: a case study based on land snails in Skyros archipelago (Aegean Sea, Greece). Journal of Biogeography 32: 1727–1735.
Welter-Schultes F. W. 2012. European non-marine molluscs, a guide for species identification. Planet Poster Edition, Göttingen.
Welter-Schultes F. W., Williams M. R. 1999. History, island area and habitat availability determine land snail species richness of Aegean islands. Journal of Biogeography 26: 239–249.
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